Translucent Tree is centered around divorced Chigiri, a suburban middle-aged mother taking care of her teenage daughter, Mayu, and her ailing father Kaho. Her life is mundane and rather hopeless until Go, a TV producer from Tokyo, pays a visit to her town and the tree they shared time under some twenty years back after a TV commercial shoot. His remembrances of her, the tree, and the long unrequited love stir both of their hearts to fumbling action as they re-discover the love between them.

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